TYPE: Costa Rica. Puntarenas: Coto Brus, Las Alturas de Cotón, Zona Protectora Las Tablas, Las Alturas Biological Station, southem slopes of Cerro Chai, 8°56'59"N 82°50'01"W, 1500 m, lower montane wet forest, primary vegetation, 26 October 2005, F. Pupulin 5839, D. Bogarín, R.L. Dressler, R. Gómez, A. Karremans, A. Rambelli & S. Rambelli (holotype, USJ; illustration of type; photo of the flower from the plant that served as the holotype).